What powers pnas.org?
Last seen Jun 2026 · 38 technologies across 19 categories
Analytics
Google Analytics
since Jun 2018
New Relic
since Aug 2018
Caching
Varnish
since Apr 2018
CDN
CMS
Databases
Ecommerce
Cart Functionality
since Aug 2020
Font scripts
Google Font API
since Aug 2018
JavaScript Libraries
Miscellaneous
Operating Systems
CentOS
since Aug 2018
PaaS
Pantheon
since May 2019
Programming languages
Retargeting
Google Remarketing Tag
since Jul 2021
RUM
Cloudflare Browser Insights
since Mar 2022
Security
Cloudflare Bot Management
since Mar 2022
HSTS
since Sep 2022
Tag managers
Google Tag Manager
since Jan 2019
UI frameworks
Web Servers
Nginx
since Aug 2018
Top search keywords
| Keyword | Position | Est. monthly visits |
|---|---|---|
| sexes | #7 | 46,800 |
| cystathionine beta synthase | #7 | 17,200 |
| genetic modification | #95 | 4,700 |
| endoplasmic reticulum | #9 | 4,500 |
| pnas | #1 | 4,500 |
| proc natl acad sci u s a | #1 | 4,500 |
| wolf and dog | #3 | 4,400 |
| sexes | #71 | 3,800 |
| prions | #4 | 3,000 |
| functional dna | #1 | 2,500 |
| phylogenetic history | #5 | 1,900 |
| science and social science | #4 | 1,800 |
| equitability | #10 | 1,500 |
| glp-1 | #25 | 1,400 |
| cystathionine beta synthase | #51 | 1,400 |
| cystathionine beta synthase | #25 | 1,400 |
| academy | #26 | 1,400 |
| atmospheric oxygen | #3 | 1,300 |
| neanderthal | #15 | 1,300 |
| how many cells in the human body | #2 | 1,200 |
| echo chamber | #4 | 1,200 |
| sister chromatids | #5 | 1,000 |
| dmt | #12 | 1,000 |
| daylight time change | #59 | 870 |
| cod fisheries | #11 | 820 |
The site's top keywords in Google (US), with the visits each one is estimated to bring per month.
Top backlinks
Top links pointing at the site, one per source domain, strongest first. The anchor text opens the page carrying the link.
Audience
- 82% desktop
- 17% phone
- 1% tablet
United States
India
Japan
Germany
United Kingdom
France
Countries where the site has Chrome traffic, strongest markets first.
Open Page Rank
8.91 / 10 · ranked #528 of all domains
33,516 referring domains · OpenPageRank
Search traffic
398,700 est. monthly organic visits (US)
240,127 keywords in Google's top 30
Core Web Vitals
| LCP | 2.8 s |
| INP | 100 ms |
| CLS | 0.05 |
| TTFB | 1500 ms |
Real user p75, Chrome UX Report, Jun 2026.
Page
- Language: English
- CDN: Cloudflare
- Weight: 1.8 MB · 45 requests
- Structured data: JSON-LD, OpenGraph
Homepage as crawled by the HTTP Archive, Jul 2026.
Subdomains
| Host | Techs |
|---|---|
| blog.pnas.org | 27 |
| www.pnas.org | 17 |
| chemistry.pnas.org | 10 |
| frontmatter.pnas.org | 10 |
| sustainability.pnas.org | 10 |
| physics.pnas.org | 7 |